Proceedings of the Biological Society of Washington
Although the mites belonging to the family Chaetodactylidae are commonly found on bees of the genus Osmia, and related genera, little is known about them. Trouessart (1904), working with Chaetodactylus ludwigi (Trouessart), and Popo-vici-Baznosanu (1913), working with C. osmiae (Dufour), discovered the encysted hypopial stage. They believed that this was an overwintering form that transformed into an adult in the spring. Michael (1903) reared C. osmiae from the migratory hypopus through the third nymph into the adult. However, he did not find the encysted hypopial stage. Zakhvatkin (1941) has presented the most complete bibliography of the family Chaetodactylidae.
Baker, Edward W., "XV. Descriptions of the Stages of Chaetodactylus Krombeini, New Species, a Mite Associated with a Bee, Osmia Lignaria Say (Acarina: Chaetodactylus)" (1962). An. Paper 253.
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