Aggressive Behavior in a Centrolenid Frog, With Comments on Territoriality in Anurans
Centrolenella griffithsi (Goin) ♂ ♂ exhibit aggressive behavior consisting of grasping one another with the forelimbs while suspended from vegetation. Aggressive behavior and territoriality are well developed in certain species having specialized reproductive modes that require limited resources.
Duellman, William E., and Alan H. Savitzky. 1976. Aggressive behavior in a centrolenid frog, with comments on territoriality in anurans. Herpetologica, 32(4):401-404.